Wednesday, April 2, 2008

Fruit development


A fruit is a ripened ovary. After the ovule in an ovary is fertilized in a process known as pollination, the ovary begins to ripen. The ovule develops into a seed and the ovary wall pericarp may become fleshy (as in berries or drupes), or form a hard outer covering (as in nuts). In some cases, the sepals, petals and/or stamens and style of the flower fall off. Fruit development continues until the seeds have matured. With some multiseeded fruits the extent to which the flesh develops is proportional to the number of fertilized ovules.
The wall of the fruit, developed from the ovary wall of the flower, is called the pericarp. The pericarp is often differentiated into two or three distinct layers called the exocarp (outer layer - also called epicarp), mesocarp (middle layer), and endocarp (inner layer). In some fruits, especially simple fruits derived from an inferior ovary, other parts of the flower (such as the floral tube, including the petals, sepals, and stamens), fuse with the ovary and ripen with it. The plant hormone ethylene causes ripening. When such other floral parts are a significant part of the fruit, it is called an accessory fruit. Since other parts of the flower may contribute to the structure of the fruit, it is important to study flower structure to understand how a particular fruit forms.
Fruits are so varied in form and development, that it is difficult to devise a classification scheme that includes all known fruits. Many common terms for seeds and fruit are incorrectly applied, a fact that complicates understanding of the terminology. Seeds are ripened ovules; fruits are the ripened ovaries or carpels that contain the seeds. To these two basic definitions can be added the clarification that in botanical terminology, a nut is not a type of fruit and not another term for seed, on the contrary to common terminology.